Suborder Trogiomorpha Suborder Troctomorpha Suborder Psocomorpha

Infraorder Atropetae

Family LEPIDOPSOCIDAE

Subfamily Thylacellinae

Thylacella cubana (Banks) 1941

Thylacella cubana map.

Thylacella cubana female. Size: 2.5 mm Range: This is the only species of the genus in the Western Hemisphere. It has been colleted around Brownsville, TX and penninsular Florida. Outside the U.S. it occurs in Cuba, Mexico, Guatemala and Belize. Habitat: persistant dead leaves of various plants, also on trunks and branches of trees.

Subfamily Lepidopsocinae

Echmepteryx hageni (Packard) 1870

Echmepteryx hageni map

echmepteryx hageni female Size: ~ 3 mm Range outside Texas: Eastern U.S. west to Arkansas Habitat: Bark and leaves of deciduous trees and on exposed outcrops. Males are rare except at a few sites in the U.S.

Subfamily Echinopsocinae

Neolepolepis occidentalis (Mockford) 1955

Neolepolepis occidentalis map.

Neolepolepis occidentalis female Size:~1.6 mm. Range outside Texas: Eastern U.S.; less common in the southwest. Habitat: Leaf litter. There is a macropterous form of this species and I would be delighted to encounter it.

Family TROGIIDAE

Cerobasis guestfalica (Kolbe) 1880

Cerobasis guestfalica map

Cerobasis guestfalica female Size: 1.5 mm. Cosmopolitan. Habitat: Deciduous and evergreen trees and stone outcrops. Identified by the anchor shaped mark on the frons. Most populations are parthenogenetic.

Lepinotus reticulatus Enderlein 1905

Lepinotus reticulatus map

Lepinotus reticulatus female Size: ~ 1.2 mm. Range: Cosmopolitan. Habitat: Leaf litter, granaries, bird and mammal nests. Winglets with a distinctive reticulate pattern. The wings are easily dislodged. This obligate parthenogen can become a pest in stored food products.

Trogium pulsatorium (Linnaeus) 1758

Trogium pulsatorium map

Trogium pulsatorium Size: ~ 1.7 mm. Range:  Not common; found primarily in the northeast; Habitat: Can occur in grain mills, bird nests, even herbaria. I have only found it on palms (Sabal minor) at Aquarena Springs. Males and females look very much alike.

Family PSOQUILLIDAE

Rhyopsocus bentonea Sommerman 1956

Rhyopsocus bentonea map

Rhyopsocus bentonea male 1.03 mm, female 1.18 mm Range: throughout penninsular Florida, Atlantic Coast to Brunswick GA, around Gulf Coast to Texas, southern and southeastern Mexico. Habitat: dead leaves of palm,yucca and Typha. In this photo the male is on the left. I call this one the "Hobbit psocid".

Rhyopsocus eclipticus Hagen 1876

Rhyopsocus eclipticus map

Rhyopsocus eclipticus female Size: 1.4 mm. Range: Atlantic Coastal Plain, Gulf Coast from Florida to south Texas, along Mississippi embankment to southern Illinois. Habitat: dead persistent leaves, conifer foilage and leaf litter. Note the large terminal segment of the maxillary palps.

Family PSYLLIPSOCIDAE

Dorypteryx domestica (Smithers) 1958

Dorypteryx domestica map

Dorypteryx domestica Size: ~ 2 mm. Range outside Texas: records from central Europe, southern England, Zimbabwe. It is only known from domestic situations such as basements. This rather odd lookng psocid reminds me of Gollum from Lord of the Rings.

Psyllipsocus apache Mockford 2011

Psyllipsocus apache map

Psyllipsocus apache female Psyllipsocus apache male Size: 1.2 - 1.5 mm. Range outside Texas: Records from Arizona and California. Habitat: Found in leaf litter, dead yucca leaves and dead palm leaves. All are brachypterous. Named for Indian tribe from type locality.

Psyllipsocus ramburii Selys-Longchamps 1872

Psyllipsocus ramburii map

Psyllipsocus ramburii female Psyllipsocus ramburii female Size: ~2 mm Range: cosmopolitan. Females micropterous or macro- pterous. Found in caves, cellars and occasionally shaded outcrops. Records:Texas Memorial Museum.

Micropterous image by Matthew Bergeron; macropterous one by Stepen Luk.

Suborder Troctomorphareturn to top

Infraorder Amphientometae

Family AMPHIENTOMIDAE

Lithoseopsis hellmani (Mockford & Gurney) 1956

Lithoseopsis hellmani map

Lithoseopsis hellmani female Size: 3.5 mm Range outide Texas: southern Arizona south to Central America. - Habitat: Shaded limestone outcrops. This species is part of a complex of closely related species found from southern Texas. Known from female only.

Infraorder Nanopsocetae

Family LIPOSCELIDIDAE

Subfamily Embidopsocinae

Belaphotroctes sp.

Belaphotroctes female Yes, there are liposcelidids with wings! In this genus mx4 is much wider than mx3. Females may be apteous or macropterous depending on species.

Size:1.7 mm Range: Winged females have been found in Hays Co. This specimen was caught sweeping grass. This one is a possible ant mimic. We hope to find apterous males and females of this species.

Embidopsocus laticeps Mockford 1963

Embidopsocus laticeps Size: ~ 1.0 mm Range: Texas- Harris Co.; Gulf Coast Florida to Texas and in Bullock Co. Ga. Habitat: under bark of dead trees. Known from males, macropterors and apterous females.

Photoghaph by Graham Montgomery

Subfamily Liposcelidinae

Liposcelis Section I

Liposcelis brunnea Motschulsky 1852

brunnea Size: ~ 1.1 mm Range: west Texas, Arizona, New Mexico, Colorado, northeast toOttowa, Ontario. Habitat: Conifer and oak foilage, and ground litter beneath these trees, bird nests and domestic situations.

Image by George Opit.

Liposcelis deltachi Sommerman 1957

Liposcelis deltachi Size: 1.1 mm Range: Hays Co.; Southern and western Texas, southern New Mexico, southern Oklahoma; San Luis Potosi, Mexico. Habitat: Foliage of junipers, pine, yuccas, mesquite and leaf litter. Known from females only.

Liposcelis entomophila (Enderlein) 1907

Liposcelis entomophila Size: ~ 1.1 mm. Range: Texas-Hays Co.;  Midwest and southeastern U.S. to central Texas; fairly cosmopolitan. Habitat: Primarily domestic but also taken in leaf litter. Field collected specimens (as in leaf litter) mostly tropical and subtropical. Known from male and female.

Liposcelis hirsutoide Mockford 1978

Liposcelis hirsutoide Size: 1.1 mm. Range: central and south Texas and in central-penninsular Florida.  Habitat: woody vegetation and under bark. I find it especially on lichen covered banches of Mesquite.  Known from both sexes but I have yet to photograph a male.

Liposcelis ornata Mockford 1978

Liposcelis ornata female Size: 1.2 mm. Range: Gulf Coast from Florida to Texas and in the Mexican states of San Luis Potosi and Tabasco. Known from female only. Habitat: Great variety of trees and shrubs. I have found in in leaf litter and dead palm fronds.

Liposcelis pearmani Lienhard 1990

Liposcelis pearmani Size: ~ 1.1 mm. Range: Texas- Hays Co.  Relatively cosmopolitan; recorded from several U. S. states including Texas. Habitat: Domestic; also found in bird nests, under bark and sweeping grass. This species is known from both sexes.

Liposcelis pallida Mockford 1978

Liposcelis pallida Size: ~ 1 mm. Range: Texas- Blanco Co.; Known from the Davis Mts. of Texas and Catalina Mts. of Arizona. The specimen pictured was collected at Flat Creek (Blanco Co.). Habitat: Dead persistent leaves of yucca; leaf litter. Known from both sexes.

Liposcelis Section II

Liposcelis bostrychophila Badonnel 1931

Liposcelis bostrychophila female Size: ~ 1.0 mm Range: Texas- Hays Co.;  Cosmopolitan Habitat: under bark, bird nests, stored grain, old books, chicken litter. Known only from females. If there is a classic booklouse, this is it.

Family PACHYTROCTIDAE

Nanopsocus oceanicus Pearman 1928

Nanopsocus oceanicus map

Nanopsocus oceanicus female Nanopsocus oceanicus male Size: ~ macropterous female , 1.5 mm., apteous female 1.0 mm. Range: southeastern U. S. north to Illinois; also Mexico, Central America, West Indies, West and Central Africa, Japan and South Pacific Islands . Habitat: Occurs on dead leaves of palms, bamboos and grasses.

Nanopsocus oceanicus mac female

Nanopsocus oceanicus mac female Nanopsocus oceanicus mac female Macropterous females brown;apterous females creamy white; males (apterous) creamy white but with a reddish brown stripe along anterior margin of abdominal terga 2-7. This genus is characterized by a foliate shaped hind tarsal claw on each foot.

Peritroctes bengalensis Thornton 1966

Peritroctes bengalensis map

Peritroctes bengalensis female Size: 1.1 mm. Range: First recorded from the Botanical Gardens of Calcutta, India by Ian W. B. Thornton and S. K. Wong, "Some Psocoptera from West Bengal, India", Transactions of the Royal Entomological Society of London, 118(1):1-21, 1966. Habitat: leaf litter, dead palm frinds, dead persistent leaves.

I have a colony in my leaf litter/ dead palm frond habitat here in San Marcos, TX. I am not sure what the big picture is.

Tapinella maculata Mockford and Gurney 1956

Tapinella maculata map

Tapinella maculata male Tapinella maculata female Size: male ~ 1 mm, female ~ 1.5 mm. Range: Central Texas to lower Rio Grande Valley, Mexico and South America. Habitat: dead palm and grass leaves Known from males, macropterous and apterous females.

Tapinella maculata female Tapinella maculata male To the left is the macropterous female (length 1.7 mm). In my experience, this form is rarer than the apterous one. In this family, the wings are held flat over the abdomen, not roof-like as in the Psocomorpha. Dr. Mockford says that this individual from Gonzales Co. shows much more patterning on the head than specimens in far south Texas.

Tapinella sp. "Type 2"

Tapinella sp. map

Tapinella sp. female Tapinella sp. male Size: male ~ 1.1 mm, female ~ 1.3 mm Range: unknown Habitat: Leaf litter This un-described species lacks the backwards "U" shaped marks along the lateral edge of the abdomen. This species often occurs with Tapinella maculata.

Tapinella sp. Type 2 Tapinella sp. Type 2 The images on this row depict the macropterous form of Tapinella sp. Type 2. These individuals possess ocelli while the apterous morphs do not.

Family SPHAEROPSOCIDAE

Sphaeropsocus bicolor Mockford 2013

Sphaeropsocus bicolor map

Sphaeropsocus bicolor Size: 0.7 mm Range: single site Habitat: leaf litter This species has not been found in Texas but it is so remarkable that I had to include it. "It is the first living species of its genus, known otherwise only from Sphaeropsocus kuenowii Hagen, a fossil species from the Baltic amber (Eocene)." Mockford, E.L., A New species of Sphaeropsocus Hagen from south eastern U. S.: the first living species of its genus, Life: the Excitement of Biology 1(2): 100-111.

Suborder Psocomorphareturn to top

Epipsocetae   Caeciliusetae   Homilopsocidea   Psocetae

Infraorder Epipsocetae

Family EPIPSOCIDAE

Bertkauia crosbyana Chapman 1930

Bertkauia crosbyana map

Bertkauia crosbyana female Size: ~ 1.7 mm  Range: throughout eastern U.S., southern Rocky Mts., northwest Washington state. Mexico: San Cristobal and Chiapas. Habitat: woodland ground litter Males (very rare) macropterous, femaes micropterous.

Infraorder Caeciliusetae

Caeciliusidae   Stenopsocidae   Amphipsocidae   Dasydemellidae

Family CAECILIUSIDAE

Stenocaecilius casarum (Badonnel) 1931

Stenocaecilius casarum map

Stenocaecilius casarum female Size: ~ 3.5 mm Range: outside Texas- Gulf Coast from south Florida to the Rio Grande Valley. Wide range in the tropics including Mexico, Guatemala & Venezuela. Habitat: Living or dead palm foilage. With the exception of a single male taken in Guyana, this species consists only of females.

Valenzuela croesus (Chapman) 1930

Valenzuela croesus map

Valenzuela croesus female Valenzuela croesus male Size: ~ 3 Range: Texas, Arkansas east to Mississippi, North Carolina and north the New York; ranges from Mexico to Guatemala. Habitat: Prefers foilage of Juniperus and Cupressus.

Valenzuela flavidus (Stephens) 1836

Valenzuela flavidus

Valenzuela flavidus female Valenzuela flavidus male Size: 3.6 mm, Range outside Texas: eastern U. S., northern Pacific Coast, also Europe, Canary Islands, Bermuda, and Mexico. Habitat: Foilage of broadleaf trees. Males are scarce throughout most of the range. Populations which have males may represent a sibling species (Mockford, 1993).

Valenzuela indicator Mockford 1969

Valenzuela indicator map

Valenzuela indicator female Valenzuela indicator male Size:~ 4 mm, Range outside Texas: Florida, Georgia, Cuba, Habitat: found on various trees including oaks, palms and on Sapnish moss.

Images by Robert Zimlich.

Valenzuela lochloosae (Mockford) 1965

Valenzuela lochloosae map

Valenzuela lochloosae female Valenzuela lochloosae male Size: ~ 3 mm Range: Throughout Florida and southeast Alabama. Range probably along entire Gulf Coast. Habitat: persistent dead leaves of native grasses.

Valenzuela manteri (Sommerman) 1943

Valenzuela manteri map

Valenzuela manteri female Valenzuela manteri female Size: ~ 3 mm Range outside Texas: eastern U. S. records in Minnesota, Missouri and Nebraska. Habitat: dead leaves of cattail, corn and palmetto. This species was described by Kathryn M. Sommerman in "Description and bionomics of Caecilius manteri", Proc. Ent. Soc. of Washington, 45(2): 9-39, 1943 with marvelous illustrations. Males are rare in this species.

Valenzuela micanopi (Mockford) 1965

Valenzuela micanopi map

Valenzuela micanopi female Valenzuela micanopi male Size: male 3.0 mm, female 3.2 mm Range outside Texas: throughout Florida, southeast Alabama and on Bimini Island and the Bahamas. Habitat: dead leaves of Sabal and Coccothrinax.

Valenzuela nadleri Mockford 1966

Valenzuela nadleri map

Valenzuela nadleri female Valenzuela nadleri male Size: male 2.6 mm, female 2.7 mm. Range: eastern US, west to Wisconsin and Missouri,also SE Canada. Recent records extend range westward. Habitat: leaf litter. Females dimorphic- macropterous and micropterous.

Valenzuela nadleri micro female This is an image of the micropterous female by Jean Brodeur of Canada. This female morph looks very similar to the micropterous female of Valenzuela posticus. Micropterous females of V. nadleri, however, have dark antennae all the way to the tip. Both  occur in leaf litte

Valenzuela subflavus (Aaron) 1886

Valenzuela
subflavus map

Valenzuela subflavus female Valenzuela subflavus male Size: ~ 2.5 mm  Range: Outside Texas - Atlantic and Gulf Coast  Habitat:  foliage of broadleaf evergreen trees like American holly and live oak.

Family STENOPSOCIDAE

Graphopsocus cruciatus (Linnaeus) 1768

Graphopsocus cruciatus map

Graphopsocus cruciatus female Graphopsocus cruciatus male Size:  ~3.8 mm   Range:  Texas Gulf Coast. This non-native species was apparently introduced along both coasts sometime in the early 19th century. Habitat: Foilage of braodleaf deciduous and evergreen trees.

Family AMPHIPSOCIDAE

Polypsocus corruptus (Hagen) 1861

Polypsocus corruptus map

Polypsocus corruptus female Polypsocus corruptus male Size: ~ 3.5 mm  Range outside Texas: eastern U. S. west to Missouri; eastern Canada and Pacific coast from British Columbia south to northern Califorina. There are closely related species in Mexico. Habitat: Variety of broad-leaf  trees including Magnolia.

Family DASYDEMELLIDAE

Teliapsocus conterminus (Walsh) 1863

Teliapsocus conterminus

Teliapsocus conterminus female Teliapsocus conterminus male Size: 5.6 mm Range: Throughout most of U.S. and Canada, but absent from northern midwestern states. Habitat: Great variety of broad-leaf and coniferous trees and occasionally ground litter. Not known outside North America.

Male image by Patrick Coin.

Infraorder Homilopsocidea

Lachesillidae  Ectopsocidae  Peripsocidae  Trichopsocidae  Archipsocidae  Pseudocaeciliidae

Family LACHESILLIDAE

Andra Group

Lachesilla kola Sommerman 1946

Lachesilla
kola map

Lachesilla ola female Lachesilla kola male Size: ~ 2.2 mm  Range: Outside Texas- Records from southern California, Arizona and New Mexico  Habitat: specimens shown found on dead Chinaberry branches near the Concho River.

Lachesilla nubilis (Aaron) 1886

Lachesilla nubilis map

Lachesilla nubilis female Lachesilla nubilis male Size: ~ 2.6 Range: Outside Texas- throughout eastern U.S. west to New Mexico, Colorado and Utah.  Habitat: dead grasses and dead limbs with persistent leaves.

Lachesilla texana Mockford & Garcia Aldrete 2010

Lachesilla texana map

Lachesilla texana female Lachesilla texana male Size: ~2.2 Range: Outside Texas - a second population in the Smokey Mts. Mexico: Nuevo Leon, Coahuila, and Cerrode la Silla. Habitat: dead leaves on clumps of native grasses.

Corona Group

Lachesilla michiliensis Garcia Aldrete 1991

Lachesilla michiliensis map

Lachesilla michiliensis female Size: ~ 1.5 mm Range: Outside Texas  Mexico: Durango State Habitat: under loose bark Known only from female. "This species is neotenic as indicated by the brachyptery, small oceli and almost complete absence of sensory fields on the paraprocts" Garcia Aldrete, A., "Lachesillidae from the Biosphere Reserve 'La Michilia", Durango and Surrounding Areas, Folia Entomologica Mexicana 81: 165-183, 1991.

photograph by Graham Montgomery

Forcepeta Group

Lachesilla anna Sommerman 1946

Lachesilla anna map

Lachesilla anna female Lachesilla anna male Size: 3.2 mm Range: Outside Texas occurs throughout eastern U.S. and southeastern Canada, south to Marian Co. Florida and west to Texas Co. Missouri. Habitat: persistent dead leaves trees and shrubs.

Lachesilla contraforcepeta Chapman 1930

Lachesilla contraforcepeta map

Lachesilla contraforcepeta female Lachesilla contraforcepeta male Size: ~ 2.0 Range: Outside Texas- throughout eastern U.S., and Rocky Mountain region, New Mexico, Colorado, Idaho. Habitat: conifers such as junipers and pine. Male claspers curve outward.

Lachesilla floridana Garcia Aldrete 1999

Lachesilla floridana map

Lachesilla floridana female Lachesilla floridana male Size: 2.0 mm  Range: Outside Texas coastal distribution in Florida and Alabama;Habitat: Found on dead palm and palmetto fronds. Note the crossvein between the pterostigma and vein Rs.

Lachesilla forcepeta Chapman 1930

Lachesilla forcepeta map

Lachesilla forcepeta female Lachesilla forcepeta male Size: ~ 2.0 mm Range: Outside of Texas- throughout eastern U.S. west to Oklahoma and Kansas and the San Antonio area of Texas. Habitat: conifers including Juniperus ashei.

Lachesilla forcepeta Chapman 1930

Lachesilla forcepeta map

Lachesilla forcepeta female Lachesilla forcepeta male Size: ~ 2.0 mm Range: Outside of Texas- throughout eastern U.S. west to Oklahoma and Kansas and the San Antonio area of Texas. Habitat: conifers including Juniperus ashei.

Lachesilla kathrynae Mockford & Gurney 1956

Lachesilla kathrynae map

Lachesilla kathrynae female Lachesilla kathrynae male Size: ~ 2.5 mm Range: Lower Rio Grande Valley; Mexico: Tabasco State Habitat: dead foliage of palms and carrizo grass.

Lachesilla penta Sommerman 1946

Lachesilla penta map

Lachesilla penta female Lachesilla penta male Size: ~ 2.5 Range: Outside Texas- Gulf states from Texas to Florida; eastern Mexico south to Chiapas State. Habitat: prsistant dead leaves of deciduous trees and palms.

Pedicularia Group

Lachesilla pedicularia (Linnaeus) 1758

Lachesilla pedicularia map

Lachesilla pedicularia female Awaiting image of male Size: 2.5 mm. Range: This species has been recorded from most of the U.S., nearly cosmopolitan elsewhere. Habitat: Found on grasses, dead persistent leaves, dried grain and occcasionally conifers. It is easily transported in human commerce and can become a pest of stored grain.

Lachesilla rena Sommerman 1946

Lachesilla rena map

Lachesilla rena female Awaiting image of male Size:1.8; Range: Outside Texas - southern California and Maricopa Co. Arizona and throughout much of Mexico. Habitat: dead grasses and low vegetation.

Lachesilla tectorum Badonnel 1931

Lachesilla tectorum map

Lachesilla tectorum female Size: 2.5 mm Range: outside Texas- Gulf coast  Florida to Texas Mexico: Tamaulipas and Nuevo Leon Habitat: dead leaves of palm and tall grasses Known from female only.

Lachesilla yucateca Mockford 2002

Lachesilla yucateca map

Lachesilla yucateca female Lachesilla yucateca male Size: ~ 1.7 mm Range: Outside Texas- Mexico: Oaxaca, Tamaulipas, Yucatan; Habitat: persistent dried leaves of native grasses. This species belongs to the Rena complex in which the male epiproct is extremely long and narrow.

Riegeli Group

Lachesilla tropica Garcia Aldrete 1982

Lachesilla tropica map

Lachesilla tropica female Lachesilla tropica male Size: 2.2 mm Range - Outside Texas: Florida, throughout most of Mexico, Guatemala, Honduras, Jamaica. Habitat: hanging dead leaves of trees and shrubs.

Rufa Group

Lachesilla nita Sommerman 1946

Lachesilla nita map

Lachesilla nita female Lachesilla nita male Size: 2.5 mm. Range: Gulf Coast Georgia to Texas, also in Mexico, Beliize, Guatemala, and Panama. Habitat: Found on cabbage palm, various oaks, Eleagnus and other shrubs. Note the characteristic diffuse spot in each cell from R3, R5, M1, M2, M3.

Sclera Group

Lachesilla sulcata Garcia Aldrete 1986

Lachesilla sulcata map

Lachesilla sulcata female Size: ~ 2.0 mm Range: Outside Texas- Gulf states from Texas to Florida. Mexico: Nuevo Leon, Tamaulipas, Chiapas. Habitat: native grass.The male of this species was only recently been described (Garcia Aldrete & Mockford "Reappraisal of Species Group Patzunensis" Proc. Ent. Soc. Washington 113(4): 417-425, 2011.

Family ECTOPSOCIDAE

Ectopsocopsis cryptomeriae (Enderlein) 1907

Ectopsocopsis cryptomeriae map

Ectopsocopsis cryptomeriae female Ectopsocopsis cryptomeriae male Size: ~ 1.7 mm Range: Outside Texas- Gulf States from Texas to Florida and north to New York, west to Illinois Habitat: leaf litter; hanging dead leaves.

Ectopsocus californicus (Banks) 1903

Ectopsocus californicus map

Ectopsocus californicus female Ectopsocus californicus male Size: ~ 2.4 Range: Outside Texas: U. S. Atlantic and Pacific Coast, Mexico and the highlands of Guatemala Habitat: living and dead foliage of broadleaf and coniferous trees.

Ectopsocus meridionalis Ribaga 1904

Ectopsocus meridionalis map

Ectopsocus meridionalis Size: 2.4 mm Range outsie Texas: throughout eastern U.S. south to Florida; central Mexico to Chile, southern Europe, Africa and Japan. Habitat: persistent dead leaves. Known only from female.

Ectopsocus vachoni Badonnel 1945

Ectopsocus vachoni map

Ectopsocus vachoni female Ectopsocus vachoni female Size: males ~ 1.0 mm, females ~ 1.7 mm Range: Outside Texas-US Gulf States,California; Mexico, South America, and Europe Habitat: leaf litter, dried grass and lower foliage of trees.

Ectopsocus vachoni female In this species, the males are micropterous and the females are either macropterous or micropterous. The tiny male has a dark clunial comb (visible in a dissecting microscope) which identifies it as mature. I once observed a male approaching a female and vibrating his body rhythmically as a prelude to courtship. This species is likely very common throughout Texas.

Family PERIPSOCIDAE

Peripsocus madidus (Hagen) 1861

Peripsocus madidus map

Peripsocus madidus female Peripsocus madidus male Size: ~3.2 mm Range: Outside Texas, Nova Scotia south to Florida and west to central Texas Habitat: bark and branches of broad-leaf and coniferous trees.

Peripsocus madidus brachypeterous The barchypterous form of this species is shown to the left. I do not find this form in central Texas but did record them from Austin Co.

Peripsocus subfasciatus (Rambur) 1842

Peripsocus subfasciatus

Peripsocus subfasciatus Size: ~ 2.7 Range: Outside Texas - eastern U.S., west to Minnesota and Arkansas, Pacific Coast from San Francisco Bay to British Columbia. Habitat: branches of broadleaf and coniferous trees and shaded stone outcrops. Male virtually absent except in Washington state and Vancouver, British Columbia.

Peripsocus minimus Mockford 1971

Peripsocus minimus map

Peripsocus minimusfemale Peripsocus minimus male Size: ~ 2.0 mm  Range:  Outside Texas- southern Illinois southwestward to Missoiri, Arkansas and central Texas  Habitat: foliage of Juniperus virginiana and Juniperus ashei.

Peripsocus sp.

Peripsocus sp. mystery map

Peripsocus sp female Peripsocus close to stagnivagus male Size: male: ~ 2.2 mm, female ~ 1.5 mm Range: Habitat: Lichen covered branches of Quercus virginiana and Texas persimmon (Diospyros texana). This is an un-described species in which the female is micropterous.

Family TRICHOPSOCIDAE

Trichopsocus dalii (McLachlan) 1867

Trichopsocus dalii map

Trichopsocus dalii female Trichopsocus dalii male Size: ~ 2.8 mm Range: outside Texas- this introduced species is found in states on the Gulf and Atlantic coast of the U.S. and in the Atlantic and Mediteranean coast of Europe and North Africa Habitat: branches of braod leaf trees like citrus, bay, Ligustrum and Ilex.

Family ARCHIPSOCIDAE

Archipsocus sp.

Archipsocus sp. Archipsocus specimens are found along the Gulf coast from Florida to Texas and in Mexico. They can occur in Texas as far inland as Travis Co. I have not yet identified this species. These specimens were found at Palmetto State Park. There are three described species which might occur on the Texas coast: Archipsocus floridanus, A. gurneyi A. nomas. The webbing from these psocids often covers entire tree trunks and causes a media sensation.

Family PSEUDOCAECILIIDAE

Pseudocaecilius citricola (Ashmead) 1879

Pseudocaecilius citricola map

Pseudocaecilius citricola female Size: 2.8 mm Range: Outside Texas, occurs throughout Florida, on the Alabama coast, pan-tropical; is even recorded on the Galapagos Islands; Habitat: living leaves of citrus, evergreen oaks, palms and other trees. Known from female only.

Family PHILOTARSIDAE

Aaroniella sp.

Aaroniella sp. map

Aaroniella sp. female Aaroniella sp. male Size: 2.2 mm This un-described species is found at the Aransas National Wildlife Refuge. Habitat: small dead branches of live oak in a heavily wooded area.

Infraorder Psocetae

Family HEMIPSOCIDAE

Hemipsocus chloroticus (Hagen) 1858

Hemipsocus chloroticus map

Hemipsocus chloroticus female Hemipsocus chloroticusHemipsocus chloroticus male Size: ~ 3 mm. Range outside Texas: Non native. Found South Carolina to Florida along coastal plain to Texas; Oregon as an introduction from Japan. Native to Southeast Asia and many Pacific Islands. Habitat: Forest leaf litter.

Family PSOCIDAE

Amphigerontiinae      Psocinae

Subfamily Amphigerontiinae

Amphigerontia montivaga (Chapman) 1930

Amphigerontia montivaga map

Amphigerontia montivaga female Amphigerontia montivaga male Size: ~ 4.7 mm Range: outside Texas- New York south to Florida and west to Wisconsin and south Texas Habitat: branches of broad-leaf and evergreen trees.

Blaste garciorum Mockford 1984

Blaste garciorum map

Blaste garciorum female Blaste garciorum male Size: ~ 3.5 mm Range outside Texas- south to Mexico and Honduras Habitat: branches of small trees and shrubs in arid woodlands.

Blaste longipennis (Banks) 1918

Blaste longipennis map

Awaiting image of female Blaste longipennis male Size: ~ 5 mm. Range outside Texas: Yellowstone Natonal Park south to Chiricahua Mountainsof Arizona. Habitat: Collected on dwarf mistletoe, Arceuthobuim vaginatum and conifers.

Blaste persimilis (Banks) 1908

Blaste persimilis map

Blaste persimilis female Blaste persimilis male Size: 3.5 mm. Range: Gulf Coast from Alabama to Brownsville, Texas then north to Refurio. Habitat: branches of oaks and dead vines. Known only from the male. I found females at Palm Harbor, TX. No published description yet of female.

Blaste posticata (Banks) 1905

Blaste posticata map

Blaste posticata female Blaste posticata male Size: ~ 3.8 mm Range outside Texas- south to Mexico, Honduras and Guatemala Habitat: broad-leaf trees, shrubs and pine.

Blaste quieta (Hagen) 1861

Blaste quieta map

Blaste quieta female Blaste quieta male Size: ~ 4.5 mm Range outside Texas- Montreal to Florida and west to South Dakota and Texas; outlier populations in Idaho and British Columbia Habitat: branches of broad-leaf and coniferous trees.

Blastopsocus lithinus (Chapman) 1930

Blastopsocus lithinus map

Blastopsocus lithinus female Awaiting image of male Size: ~ 3.7 Range outside Texas- Montreal south to Floida - west to Minnesota to Texas  Habitat: branches of broad-leaf and coniferus trees.

Blastopsocus walshi Mockford 2002

Blastopsocus walshi map

Blastopsocus walshi female Blastopsocus walshi male Size: ~ 3.3 mm Range outside Texas-Illinois south to Florida Habitat: branches of broad-leaf trees.

Subfamily Psocinae

Tribe Cerastipsocini

Cerastipsocus venosus (Burmeister) 1839

Cerastipsocus venosus map

Cerastipsocus venosus female Cerastipsocus venosus male Size: male ~ 6 mm female 8mm Range outside Texas - Maine south to southern Florida; south to Mexico. Habitat: Trunks and brances of broad leaf trees and conifers. Pterostigma of males id dark while those of females are white.

Images by Robert Zimlich.

Cerastipsocus trifasciatus (Provancher) 1876

Cerastipsocus trifasciatus map

Cerastipsocus trifasciatus female Awaiting image of male Size: ~ 6 mm. Range outside Texas: Southestern U.S. continuing through Arkansas to Arizona. Also in Mexico, Guatemala, Costa Rica and Columbia. Habitat: Tree trunks and branches and rock outcrops.

Image by William Geoghegan.

Tribe Metylophorini

Metylophorus novaescotiae (Walker) 1853

Metylophorus novaescotiae map

Metylophorus novaescotiae female Metylophorus novaescotiae male Size: 6-7 mm Range outside Texas: Nova Scotia and coastal Maine to Florida, west to Minnesota. Habitat: Branches of broad-leaf trees and shrubs. Also occurs in Mexico.

Images by Tom Murray.

Tribe Psocini

Hyalopsocus sp. close to floridanus

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Hyalopsocus #1 map

Hyalopsocus</span>
sp. female Hyalopsocus</span>
sp. female Size: male  ~5.4 mm, female ~5.6 mm Range:   Habitat: Wide variety of braodleaf and coniferous trees; usually on dead branches encrusted with lichen.

Hyalopsocus striatus (Walker) 1853

Hyalopsocus striatus #2 map

Hyalopsocus striatus 2 female Hyalopsocus striatus 2 female Size:5.3 Range outside Texas: Throughout northeastern U.S. and southeast Canada. Occurs spottily in western states. Habitat: Juniperous ashei  This species is characterized by the extensive pigment in cells Cu2 and IA in the forewing.

Psocus leidyi Aaron 1886

Psocus leidyi map

Psocus leidyi female Psocus leidyi male Size: ~ 5.4 mm Range: Maine to Minnesota, south to Florida and southwest to Arkansas with an outlier population in northern California. Habitat: Broadleaf trees, Mesquite, Hackberry, live oak.

Tribe Ptyctini

Indiopsocus bisignatus (Banks) 1904

Indiopsocus bisignatus map

Indiopsocus bisignatus female Indiopsocus bisignatus female Size: ~ 3.9 mm Range Outside Texas -eastern U.S. from northern Minnesota south to west Florida, west of the Mississippi it occurs spotily to Wyoming, Colorado and Texas; also occurs in Mexico Habitat: trunks and branches of braod-leaf and conifer trees.

Indiopsocus campestris (Aaron) 1886

Indiopsocus campestris map

Indiopsocus campestris female Indiopsocus campestris male Size: male 3.1 mm, female 3.2 mm Range: Atlantic Coast, throughout Florida continuing along Gulf Coast to Texas. Habitat: broad leaf trees and shrubs including live oak, turkey oak and pine.

Indiopsocus infumatus (Banks) 1907

Indiopsocus infumatus map

Indiopsocus infumatus female Indiopsocus infumatus male Size: ~ 3.6 Range Outside Texas - Oklahoma, New York, south to Kentucky; central Texas is probably at the western end of the range for this species   Habitat: dead branches of broad-leaf trees; my specimens on live oak and ashe juniper.

Indiopsocus lanceolatus Mockford & Young 2015

Indiopsocus lanceolatus map

Indiopsocus lanceolatus female Indiopsocus lanceolatus male Size: ~3.6 mm Range: Outside Texas: Gulf Coast, AL, GA, FL, VA. Habitat: broad leaf trees like Quercus virginiana. Recently described in Trans. Am. Entomol. Soc. 141:233-251.

Indiopsocus lacteus Mockford & Young 2015

Indiopsocus lacteus map

Indiopsocus lacteus female Indiopsocus lacteus male Size: ~ 3.9 Range: endemic to Texas Hill Country. Habitat: lower bare limbs of living Juniperus ashei. Recently described in Trans. Am. Entomol. Soc. 141:233-251.

Indiopsocus palmatus Mockford & Young 2015

Indiopsocus palmatus map

Indiopsocus palmatus female Indiopsocus palmatus male Size: ~ 3.8 Range : Apparently a Texas endemic. Habitat: braodleaf trees like Quercus virginiana, Celtis reticulata Ulmus crassifolia and Acacia farnesiana. Recently described in Trans. Am. Entomol. Soc. 141:233-251.

Indiopsocus texanus (Aaron) 1886

Indiopsocus texanus map

Indiopsocus texanus female Indiopsocus texanus male Size: ~ 3.9 mm Range outside Texas: Throughout Florida up the Atlantic coast to Virginia; Gulf Coast. Habitat: broadleaf trees including live oak.

Loensia moesta (Hagen) 1861

Loensia moesta map

Loensia moesta female Loensia moesta male Size: ~ 4.2 mm Range Outside Texas- Minnesota south to Florida west to Texas Habitat: wide variety of broad-leaf and coniferous trees.

Psocidus sp. "Beauty Queen"

Loensia beauty queen map

Loensia beauty queen female Loensia beauty queen male Size: ~ 4.3 mm Range outside Texas- unknown Habitat: lichen covered branches of live oak, Mesquite, Ashe juniper and cedar elm  In my humble opinion, this un-described species is a beautiful psocid! Don't you agree? This species is placed in a "holding" genus until further study.

Ptyctapolluta (Walsh) 1862

Ptycta polluta map

Ptycta polluta female Ptycta polluta male Size: ~ 3.4 mm Range: outside Texas- Nova Scotia south to Florida, west to Minnesota Habitat: branches of braod-leaf and coniferous trees.

Trichadenotecnum slossonae (Banks) 1903

Trichadenotecnum slossonae map

Trichadenotecnum slossonae female Trichadenotecnum slossonae male Size: ~ 3.2 mm Range: outside Texas- eastern U.S. west to Minnesta and central Texas Habitat: branches of broad-leaf and coniferous trees.

Family MYOPSOCIDAE

Lichenomima sparsa (Hagen) 1861

Lichenomima sparsa map

Lichenomima sparsa female Lichenomima sparsa male Size: male 4.7 mm female 5.9 mm Range: Outside Texas - Washington D.C. south to Florida, west to Indiana,Arkansas. Inhabits shaded stone outcrops and tree trunks. Dr. Mockford has expressed the need for this genus to be revised.

Lichenomima sp. "W"

Lichenomima sp. 'W' map

Lichenomima sp. 'W' female Awaiting image of male Size: female 4.8 mm Range: Outside Texas- unknown. Found on Quercus virginiana at Welder Wildlife Refuge.